2026
McLeod, D.S., Weisenbeck, D., Giman, B. & Chan, K.O. (2026) Amphibians and reptiles of Lanjak Entimau Wildlife Sanctuary, Sarawak, Malaysia. Zootaxa, in review
Som, H.E., Herr, M.W., Chan, K.O., Siler, C.D., Sanguila, M.B., Brown, R.M. (2026) Salinity tolerance influences biogeographic structure in island frogs: A comparative genomic study of Fejervarya in the Philippine Archipelago. In review
Pitogo, K.M.E., Meneses, C.G., Decena, S.C.P., Supsup, C.E., Som, H.E., Bernstein, J.M., Chan, K.O., Herr, M.W. & Brown, R.M. (2026) Sampling bias obscures biodiversity patterns, reveals data gaps in priority conservation areas: a call for improved documentation. PLoS One 21(7): e0353393. https://doi.org/10.1371/journal.pone.0353393
Chan, K.O. & Grismer, L.L. (2026) Extending GroupStruct2: a Bayesian and machine-learning framework for testing taxonomic hypotheses using morphometric data. ZooKeys, 1276, 125–138
Chan, K.O., Neokleous, D.N., Anuar, S., Brown, R.M., Hutter, C.R., Das, I. & Hertwig, S.T. (2026) A Genomic Perspective on Cryptic Species Reveals Complex Evolutionary Dynamics in the Gray Zone of the Speciation Continuum. Systematic Biology, syag001. https://doi.org/10.1093/sysbio/syag001
2025
Chan, K.O. & L. Lee Grismer. (2025). GroupStruct2: A User-Friendly Graphical User Interface for Statistical and Visual Support in Species Diagnosis. Systematic Biology, syaf090
DeRaad, D., DeCicco, L.H., Ostrow, E.N., Chan, K.O., Hosner, P.A., & Moyle, R.G. (2025). Rapid divergence with gene flow creates intractable nodes in the tree of life: An empirical demonstration in the Buzzing Flowerpecker (Dicaeum hypoleucum). Evolutionary Journal of the Linnean Society, 4(1), kzaf021. https://doi.org/10.1093/evolinnean/kzaf021
Torres, J., Reynolds, R. G., Chan, K. O., Hime, P., & Glor, R. E. (2025). Overwater dispersal from a single source island shaped the diversity of green anole lizards (Anolis carolinensis subgroup). Molecular Phylogenetics and Evolution, 214, 108474. https://doi.org/10.1016/j.ympev.2025.108474
Grismer, L.L., Kaatz, A., Grismer, J.L., Nguyen, E., Grergory, J.J., Wood, P.L.J., Murdoch, M.L., Anuar, S., Chan, K.O., Muin, M.A., Pawangkhanant, P., Suwannapoom, C., Poyarkov, N.A., & Quah, E.S.H. (2025). The taxonomy of Cyrtodactylus consobrinus (Peters, 1871) (Squamata, Gekkonidae) and the description of a new species from the Thai-Malay Peninsula. ZooKeys 1241, 105–137. https://doi.org/10.3897/zookeys.1241.149552
Pitogo, K.M.E, Meneses, C.G., Flores, A.B.A., Saavedra, A.J.L., Amarga, A.K.S., Angeles, M.D., Lucanas, C.C., Decena, S.C.P., Venturina, R.E.L., Fidelino, J.S., Pantinople, D., Cabanas, A.J.C., Herr, M.W, Bernstein, J.M., Chan, K.O., Sanguila, M.S., Mallari, N.A., Brown, R.M. & Supsup, C.E (2025) Caught in statistical noise: Pitfalls of a unidimensional approach to understanding biodiversity-conflict relationships. npj Biodiversity 4, 17. https://doi.org/10.1038/s44185-025-00088-4
Chan, K.O., Hime, P. & Brown, R.M. (2025). Deforestation-induced hybridization in Philippine frogs creates a distinct phenotype with an inviable genotype. Heredity 134, 200–208. https://doi.org/10.1038/s41437-025-00748-y.
2024
Kosch, T.A., Torres‑Sanchez, M.A., Liedtke, H.C., Summers, K., Yun, M.H, Crawford, A.J., Maddock, S.T., Ahammed, M.S., Arajo, V.L.N., Bertola, L.V., Bucciarell, G.M.,Carn, A., Carneiro, C.M., Chan, K.O., Chen, Y., Crottini, A., da Silva, J.M., Denton, R.D., Dittrich, C., Themudo, G.E., Farquharson, K.A., Forsdick, N.J., Gilbert, E., Che, J., Katzenback, B.A., Kotharambath, R., Levis, N.A., Marquez, R., Glib, M., Mulder, K.P., Müller, H., O’Connell, M.J., Orozco‑terWengel, P., Palomar, G., Petzold, A., Pfennig, D.W., Pfennig, K.S., Reichert, M.S., Robert, J., Scherz, M.D., Siu‑Ting, K., Snead, A.A., Stock, M., Stuckert, A.M.M., Stynoski, J.L., Tarvin, R., Valero, W. & The Amphibian Genomics Consortium (2024). The Amphibian Genomics Consortium: advancing genomic and genetic resources for amphibian research and conservation. BMC Genomics 25, 1125. https://doi.org/10.1186/s12864-024-10899-7
Chen, K.-T., Tu, Y.-J., Chan, K. O., & Teo, E. W. (2024). A Cyan-Blue Bamboo Pitviper (Trimeresurus stejnegeri stejnegeri) in Taiwan. Reptiles & Amphibians, 31, e21421.
Chan, K.O. (2024) Next-generation bioinformatics: An ultrafast and user-friendly tool for phylogenomic data exploration. Molecular Ecology Resources, e13993. https://doi.org/10.1111/1755-0998.13993
Mahony, S., Kamei, R.G., Brown, R.M. & Chan, K.O. (2024) Unnecessary splitting of genus-level clades destabilizes amphibian taxonomy. Vertebrate Zoology 74, 249–277. https://doi.org/10.3897/vz.74.e114285
2023
Chan, K.O., Anuar, S., Sankar, A., Law, I.T., Law, I.S., Shivaram, R., Ching, C., Mulcahy, D.G. & Malhotra, A. (2023) A new cryptic species of pit viper from the Ayeyarwady and Yangon regions in Myanmar (Viperidae: Trimeresurus). ZooKeys 1186, 221-234. https://doi.org/10.3897/zookeys.1186.110422
Nachman, M.W., Beckman, E.J., Bowie, R.C.K., Cicero, C., Conroy, J., Dudley, R., Hayes, T.B., Koo, M.S., Lacey, E.A., Martin, C.H., Mcguire, J.A., Patton, J.L., Spencer, C.L., Tarvin, D., Wake, M.H., Wang, I.J., Achmadi, A., A, S.T., Andersen, M.J., Arroyave, J., Austin, C.C., Barker, K., Barrow, L.N., Barrowclough, G.F., Bates, J., Bauer, A.M., Bell, K.C., Bell, R.C., Bronson, A.W., Brown, R.M., Burbrink, T., Burns, K.J., Cadena, C.D., Cannatella, D.C., Castoe, A., Chakrabarty, P., Colella, J.P., Cook, J.A., Cracraft, L., Davis, D.R., Rabosky, A.R.D., Elı, G.D., Dumbacher, P., Dunnum, J.L., Edwards, S. V, Esselstyn, J.A., Fjelds, J., Flores-villela, O.A., Ford, K., Greene, W., Hackett, S., Hamidy, A., Hanken, J., Haryoko, T., Hawkins, T.R., Heaney, L.R., Hillis, D.M., Hollingsworth, B.D., Hornsby, A.D., Hosner, P.A., Irham, M., Jansa, S., Joseph, L., Kirchman, J.J., Laduc, T.J., Jime, A., Mason, N.A., Leache, D., Mccormack, E., Mcmahan, C.D., Moyle, R.G., Ojeda, R.A., Olson, E., Chan, K.O., Parenti, L.R., Parra-olea, G., Patterson, B.D., Pauly, G.B., Pavan, S.E., Peterson, A.T., Poe, S., Rabosky, L., Raxworthy, C.J., Reddy, S., Rico-guevara, A., Rowley, J., Ruane, S., Salazar-valenzuela, D., Shultz, A.J., Streicher, W., Stuart, B.L., Summers, A.P., Tavera, J., Teta, P., Thompson, W., Timm, R.M., Torres-carvajal, O., Voelker, G., Voss, R.S., Winker, K., Witt, C., Wommack, E.A., Zink, R.M., 2023. Specimen collection is essential for modern science. PLoS Biology 21, e300231. https://doi.org/10.1371/journal.pbio.3002318
Luedtke, J.A., Chanson, J., Neam, K., Hobin, L., Maciel, A.O., Catenazzi, A., Hamidy, A., Aowphol, A., Jean, A., Sosa-Bartuano, ngel, Fong, A.G., Silva, A., Fouquet, A., Angulo, A., Kidov, A.A., Muamp, A., Saravia, oz, Diesmos, A.C., Tominaga, A., Shrestha, B., Gratwicke, B., Tjaturadi, B., Martamp, C.C., Rivera, nez, Vamp, C.R., Almazamp, squez, Seamp, C., Chandramouli, S.R., Stramp, C., Fabiola Cortez Fernamp, C., Azat, C., Hoskin, C.J., Hilton-Taylor, C., Whyte, D.L., Gower, D.J., Olson, D.H., Cisneros-Heredia, D.F., Josamp, D., Nagombi, E., Najafi-Majd, E., H Quah, E.S., Bolaamp, F., Xie, F., Brusquetti, F., amp, F.S., Andreone, F., Glaw, F., Enrique Castaamp, F., Kraus, F., Parra-Olea, G., Chaves, G., Medina-Rangel, G.F., Gonzamp, G., Mauricio Ortega-Andrade, H., Machado, F., Das, I., Ribeiro Dias, I., Nicolas Urbina-Cardona, J., Crnobrnja-Isailoviamp, J., Yang, J.-H., Jianping, J., Tshelthrim Wangyal, J., L Rowley, J.J., Measey, J., Vasudevan, K., Chan, K.O., Vasudeva Gururaja, K., Ovaska, K., Warr, L.C., Canseco-Mamp, L., Toledo, F., Damp, L.M., Monirul Khan, M.H., Meegaskumbura, M., Acevedo, M.E., Felgueiras Napoli, M., Ponce, M.A., Vaira, M., Lampo, M., Yamp, M.H., Scherz, M.D., Ramp, M.-O., Matsui, M., Fildor, M., Kusrini, M.D., Firoz Ahmed, M., Rais, M., Kouamamp, G.G., Garcamp, N., Legrand Gonwouo, N., Burrowes, P.A., Imbun, P.Y., Wagner, P., R Kok, P.J., Joglar, R.L., Auguste, R.J., Albuquerque Brandamp, R., Ibamp, R., May, R., Blair Hedges, S., Biju, S.D., Ganesh, S.R., Wren, S., Das, S., Flechas, S. V, Ashpole, S.L., Robleto-Hernamp, S.J., Loader, S.P., Inchamp, S.J., Garg, S., Phimmachak, S., Richards, S.J., Slimani, T., Osborne-Naikatini, T., F Abreu-Jardim, T.P., Condez, T.H., Carvalho, T.R., Cutajar, T.P., Pierson, T.W., Nguyen, T.Q., Kaya, ur, Yuan, Z., Long, B., Langhammer, P. & Stuart, S.N. (2023) Ongoing declines for the world’s amphibians in the face of emerging threats Threatened and extinct species. Nature, https://doi.org/10.1038/s41586-023-06578-4
Chan, K.O., Mulcahy, D.G. & Anuar, S. (2023) The Artefactual Branch Effect and Phylogenetic Conflict: Species Delimitation with Gene Flow in Mangrove Pit Vipers (Trimeresurus purpureomaculatus-erythrurus complex). Systematic Biology syad043, https://doi.org/10.1093/sysbio/syad043
Teo, E.W., Wong, S.P., Kem, Z. & Chan, K.O. (2023) Checklist of the herpetofauna of Batu Caves, Selangor, Malaysia. Malayan Nature Journal 75(1), 161–174.
Shakya, S.B., Alexander, A., Lim, H.C., Manthey, J.D., Prawiradilaga, D., Chan, K.O., Sheldon, F.H. & Moyle, R.G. (2023) Demographic History of a Complex Polymorphism in Populations of the Oriental Dwarf Kingfisher, Ceyx erithaca/rufidorsa of Southeast Asia. IBIS https://doi.org/10.1111/ibi.13207
Badli-Sham, B.H., Syafiq, M.F., Aziz, M.S.A., Jalil, N.R.M., Awang, M.T., Othman, M.N.A., Aziz, A.B.A., Dzu, K., Wahab, N.A.A., Jamil, N.L., Ismail, M.A., Azman, W.A.A.W., Ooi, X.W., Jamaha, M.A., Aqmal-Naser, M., Fahmi-Ahmad, M., Shahirah-Ibrahim, N., Rizal, S.A., Belabut, D.M., Chan, K.O., Quah, E.S.H., Grismer, L.L. & Ahmad, A.B. (2023) A decade of amphibian studies (Animalia, Amphibia) at Sekayu lowland forest, Hulu Terengganu, Peninsular Malaysia. Zookeys 1157, 43–93. https://doi.org/10.3897/zookeys.1157.95873
Chan, K.O., Grismer, L.L., Santana, F., Pinto, P., Loke, F.W. & Conaboy, N. (2023) Scratching the surface: A new species of Bent-toed Gecko (Squamata: Cyrtodactylus) from Timor-Leste of the darmandvillei group marks the potential for future discoveries. ZooKeys 1139, 107–126. https://doi.org/10.3897/zookeys.1139.96508
2022
Tan, P.E., Belabut, D., Chan, K.O. & Ahmad, N. (2022) Amphibian diversity and species composition in a lowland tropical rainforest of Ulu Gombak, Selangor, Peninsular Malaysia. Malayan Nature Journal 74(4), 521–535.
Mendoza, J.C.E., Chan, K.O., Lai, J.C.Y., Thoma, B.P., Clark, P.F., Guinot, D., Felder, D.L. & Ng, P.K.L. (2022) A comprehensive molecular phylogeny of the brachyuran crab superfamily Xanthoidea provides novel insights into its systematics and evolutionary history. Molecular Phylogenetics and Evolution 177, 107627. (Corresponding author). https://doi.org/10.1016/j.ympev.2022.107627
Chan, K.O., Law, I.S., Law, I.T., Sankar, A., Shivaram, R., Aowphol, A., Rujirawan, A., Anuar, S., Mulcahy, D., Grismer, J.G. & Grismer, L.L. (2022). Phylogeography of mangrove pit-vipers (Viperidae, Trimeresurus erythrurus-purpureomaculatus). Zoologica Scripta 61, 664–675. https://doi.org/10.1111/zsc.12562.
Sankar, A., Law, I.T., Law, I.S., Shivaram, R., Abraham, R.K. & Chan, K.O. (2022) Morphology, phylogeny, and species delimitation of Micryletta (Anura: Microhylidae) reveals a new species from Singapore. Vertebrate Zoology 72, 457–467. (Corresponding author). https://doi.org/10.3897/vz.72.e85020
Chan, K.O. & Grismer, L.L. (2022) GroupStruct: An R package for allometric size correction. Zootaxa 5124, 471–482. https://doi.org/10.11646/zootaxa.5124.4.4
Chan, K.O., Hertwig, S.T., Neokleous, D.N, Flury, J.M. & Brown, R.M. (2022) Widely used, short 16S rRNA mitochondrial gene fragments yield poor and erratic results in phylogenetic estimation and species delimitation of amphibians. BMC Ecology and Evolution 22, 37. https://doi.org/10.1186/s12862-022-01994-y
Chan, K.O., Hutter, C.R., Wood, P.L.Jr., Su, Y.C. & Brown, R.M. (2022) Gene flow increases phylogenetic structure and inflates cryptic species estimations: A case study on widespread Philippine Puddle Frogs (Occidozyga laevis). Systematic Biology 71, 40–57. https://doi.org/10.1093/sysbio/syab034.
Chan, K.O., Ahmad, N., Anuar, S., Pui, Y.M. & Das, I. (2022) Patterns and progress of Peninsular Malaysia’s amphibian research in the 21st century (2000-2020). Raffles Bulletin of Zoology 70, 80–93.
Grismer, L.L., Chan, K.O., Lovich, R.E. & Grismer, J.L. (2022) A statistical reanalysis of morphological differentiation among island night lizards (Xantusia riversiana) from the California Channel Islands. Vertebrate Zoology 72, 1–27.
2021
Sengupta S., Purakayasta J., Anandan N., Das M., Chan K.O. & Murthy C. (2021) The odd one in: re-diagnosis and phylogenetic placement of the Assam Day Gecko, Cnemaspis assamensis Das & Sengupta 2000 (Squamata: Gekkonidae). Zootaxa 5048, 581–593.
Quah, E.S.H., Badli-sham, B.H., Rahman, M.F.A., Ahmad, A. & Chan, K.O. (2021) A new record and range extension for Philautus davidlabangi (Amphibia : Rhacophoridae) from Peninsular Malaysia. Herpetology Notes 14, 1181–1186.
Chan, K.O. & Grismer, L.L. (2021) A review of reptile research in Malaysia in the 21st century. Raffles Bulletin of Zoology 69, 364–376.
Chan, K.O. & Grismer, L.L. (2021) Integrating spatial, phylogenetic, and threat assessment data from frogs and lizards to identify areas for conservation priorities in Peninsular Malaysia. Global Ecology & Conservation 28, e01650. https://doi.org/10.1016/j.gecco.2021.e01650
Chan, K.O., Schoppe, S., Rico, E.L.B. & Brown, R.M. (2021) Molecular systematic investigation of Philippine Puddle Frogs (Anura: Dicroglossidae: Occidozyga Kuhl and Van Hasselt, 1822) reveal new candidate species and a novel pattern of species dyads. Philippine Journal of Systematic Biology14(2), 1–12.
Davis, H.T., Das I., Leache, A.D., Karin, B.R., Brennan, I.G., Jackman, T., Nashriq, I., Chan, K.O. & Bauer A.M. (2021) Genetically diverse yet morphologically conserved: Hidden diversity revealed among Bornean geckos (Gekkonidae: Cyrtodactylus). Journal of Zoological Systematics and Evolutionary Research 59, 1113–1135. https://doi.org/10.1111/jzs.12470
Grismer, L.L., Wood, P.L.Jr, Poyarkov, N.A., Minh, D.L., Kraus, F., Agarwal, I., Oliver, P.M., Nguyen, S.N., Nguyen, T.Q., Suranjan, K., Welton, L.J., Stuart, B.L., Luu, V.Q., Bauer, A.M., O’Connel, K.A., Quah, E.S.H., Chan, K.O., Ziegler, T., Ngo, H., Nazarov, R.A., Aowphol, A., Chomdej, S., Suwannapoom, C., Siler, C.D., Anuar, S., Tri, N.V. & Grismer, J.L. (2021) Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia: Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71, 101–154. https://doi.org/10.3897/vz.71.e59307
Chan, K.O. & Grismer, L.L. (2021) A standardized and statistically defensible framework for quantitative morphological analyses in taxonomic studies. Zootaxa 5023, 293–300. https://doi.org/10.11646/zootaxa.5023.2.9
Rivera-Correa, M., Baldo, D., Candioti, F.V., Goyannes, V., Orrico, D., Blackburn, D.C., Castroviejo-Fisher, S., Chan, K.O., Gambale, P., Gower, D.J., Quah, E.S.H., Rowley, J.J.L., Twomey, E. & Haas, A. (2021) Amphibians in Zootaxa: 20 years documenting the global diversity of frogs, salamanders, and caecilians. Zootaxa 4979, 57–69. https://doi.org/10.11646/zootaxa.4979.1.9
Jusoh W.F.A, Ballantyne L., Chan S.H., Wong T.W., Yeo D., Nada B. & Chan K.O. (2021) Molecular systematics of the firefly genus Luciola (Coleoptera: Lampyridae: Luciolinae) with the description of a new species from Singapore. Animals 11, 687. (Corresponding author). https://doi.org/10.3390/ani11030687
Arifin, U., Chan, K.O., Smart, U., Hertwig, S.T., Smith, E.N., Iskandar, D.T. & Haas, A. (2021) Revisiting the phylogenetic predicament of the genus Huia Yang, 1991 (Amphibia: Ranidae) using molecular data and tadpole morphology. Zoological Journal of the Linnaean Society, zlaa158. https://doi.org/10.1093/zoolinnean/zlaa158
2020
Chan, K.O., Hutter, C.R., Wood, P.L.J., Grismer, L.L., Brown, R.M. (2020) Target-capture phylogenomics provide insights on gene and species tree discordances in Old World Treefrogs (Anura: Rhacophoridae). Proceedings of the Royal Society B 287, 20202102. https://doi.org/10.1098/rspb.2020.2102
Chan, K.O., Hutter, C.R., Wood, P.L., Grismer, L.L., Das, I. & Brown, R.M. (2020) Gene flow creates a mirage of cryptic species in a Southeast Asian spotted stream frog complex. Molecular Ecology 29, 3970–3987. https://doi.org/10.1111/mec.15603
Chan, K.O., Hutter, C.R., Wood, P.L., Grismer, L.L. & Brown, R.M. (2020) Larger, unfiltered datasets are more effective at resolving phylogenetic conflict: Introns, exons, and UCEs resolve ambiguities in Golden-backed frogs (Anura: Ranidae; genus Hylarana). Molecular Phylogenetics and Evolution 151, 106899. https://doi.org/10.1016/j.ympev.2020.106899
Chan, K.O. & Brown, R.M. (2020) Elucidating the drivers of genetic differentiation in Malaysian torrent frogs (Anura: Ranidae: Amolops): a landscape genomics approach. Zoological Journal of the Linnean Society 190, 65–78. https://doi.org/10.1093/zoolinnean/zlz151
Davis, H.R., Chan, K.O., Das, I., Brennan, I.G., Karin, B.R., Jackman, T.R., Brown, R.M., Iskandar, D.T., Nashriq, I., Grismer, L.L. & Bauer, A.M. (2020) Multilocus phylogeny of Bornean Bent-Toed Geckos (Gekkonidae: Cyrtodactylus) reveals hidden diversity, taxonomic disarray, and novel biogeographic patterns. Molecular Phylogenetics and Evolution 147, 106785. https://doi.org/10.1016/j.ympev.2020.106785
Jusoh, W.F.A., Ballantyne, L. & Chan, K.O. (2020) DNA-based species delimitation reveals cryptic and incipient species in synchronous flashing fireflies (Coleoptera: Lampyridae) of Southeast Asia. Biological Journal of the Linnean Society 130, 520–532. https://doi.org/10.1093/biolinnean/blaa072. (Corresponding author).
Chan, K.O., Abraham, R.K., Grismer, L.L., Brown, R.M. (2020) A systematic review of the Pulchrana picturatacomplex, with the description of a new species from Peninsular Malaysia, Sumatra, and southern Thailand. Raffles Bulletin of Zoology 68, 880–890. (doi:10.26107/RBZ-2020-0096)
Chan, K.O., Abraham, R.K., Sanguila, M.B. & Brown, R.M. (2020) Over-splitting destabilizes the taxonomy of hylaranine frogs: A response to Chandramouli et al. (2020). Zootaxa 4877, 598–600.
Chan, K.O., Muin, M.A., Badli-Sham, B.H., Fatihah-Syafiq, M., Abraham, R.K., Ahmad, A. & Zakaria, R. (2020) Identification and species delimitation of the enigmatic Marsh Frog Pulchrana rawa (Matsui, Mumpuni, and Hamidy, 2012): Second confirmed specimen and first country record for Malaysia. Journal of Herpetology 54, 282–288. https://doi.org/10.1670/19-132
Chan, K.O., Abraham, R.K. & Badli-sham, B.H. (2020) A revision of the Asian tree toad complex Rentapia hosii(Anura: Bufonidae) with the description of a new species from Peninsular Malaysia. Raffles Bulletin of Zoology68, 595–607.
Grismer, L.L., Chan, K.O., Oaks, J.R., Neang, T. & Sokun, L. (2020) A new insular species of the Cyrtodactylus intermedius (Squamata: Gekkonidae) group from Cambodia with a discussion of habitat preference and ecomorphology. Zootaxa 4830, 75–102. https://doi.org/10.11646/zootaxa.4830.1.3
Quah, E.S.H., Grismer, L.L., Lim, K.K.P., Anuar, M.S.S. & Chan, K.O. (2020) A taxonomic revision of Asthenodipsas malaccana Peters, 1864 (Squamata: Pareidae) with a description of a new species from Borneo. Zootaxa 4729, 1–24. https://doi.org/10.11646/zootaxa.4729.1.1
2019
Chan, K.O., Muin, M.A., Anuar, S., Andam, J., Razak, N. & Aziz, M.A. (2019) First checklist on the amphibians and reptiles of Mount Korbu, the second highest peak in Peninsular Malaysia. Check List 15, 1055–1069. https://doi.org/10.15560/15.6.1055
Chan, K.O. & Grismer, L.L. (2019) To split or not to split? Multilocus phylogeny and molecular species delimitation of southeast Asian toads (family: Bufonidae). BMC Evolutionary Biology 19, 95. https://doi.org/10.1186/s12862-019-1422-3
2018
Chan, K.O., Grismer, L.L. & Brown, R.M. (2018) Comprehensive multi-locus phylogeny of Old World tree frogs (Anura: Rhacophoridae) reveals taxonomic uncertainties and potential cases of over- and underestimation of species diversity. Molecular Phylogenetics and Evolution 127, 1010–1019. https://doi.org/10.1016/j.ympev.2018.07.005
Chan, K.O., Abraham, R.K., Grismer, J.L. & Grismer, L.L. (2018) Elevational size variation and two new species of torrent frogs from Peninsular Malaysia (Anura: Ranidae: Amolops Cope). Zootaxa 4434, 250–264. https://doi.org/http://dx.doi.org/10.11646/zootaxa.4434.2.2
2017
Chan, K.O., Alexander, A.M., Grismer, L.L., Su, Y.-C., Grismer, J.L., Quah, E.S.H. & Brown, R.M. (2017) Species delimitation with gene flow: a methodological comparison and population genomics approach to elucidate cryptic species boundaries in Malaysian Torrent Frogs. Molecular Ecology 26, 5435–5450. https://doi.org/10.1111/mec.14296
Chan, K.O. & Brown, R.M. (2017) Did true frogs “dispersify”? Biology Letters 13, 20170299. https://doi.org/10.1098/rsbl.2017.0299
Brown, R.M., Prue, A., Chan, K.O., Gaulke, M., Sanguila, M.B. & Siler, C.D. (2017) Taxonomic reappraisal of the northeast Mindanao stream frog, Sanguirana albotuberculata (Inger 1954), validation of Rana mearnsi, Stejneger 1905, and description of a new species from the central Philippines. Herpetological Monographs 31, 210–231. https://doi.org/10.1655/HERPMONOGRAPHS-D-16-00009.1
2016
Chan, K.O., Grismer, L.L., Zachariah, A., Brown, R.M. & Abraham, R.K. (2016) Polyphyly of Asian tree toads, genus Pedostibes Günther, 1876 (Anura: Bufonidae), and the description of a new genus from Southeast Asia. Plos One11, e0145903. https://doi.org/10.1371/journal.pone.0145903
2015
Grismer, L.L., Wood, P.L., Anuar, S., Quah, E.S.H., Muin, M.A., Chan, K.O., Sumarli, A.X. & Loredo, A.I. (2015) Repeated evolution of sympatric, palaeoendemic species in closely related, co-distributed lineages of Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) across a sky-island archipelago in Peninsular Malaysia. Zoological Journal of the Linnean Society 174, 859–876. https://doi.org/10.1111/zoj.12254
2014
Grismer, J.L., Bauer, A.M., Grismer, L.L., Thirakhupt, K., Aowphol, A., Oaks, J.R., Wood, P.L., Chan, K.O., Thy, N., Cota, M. & Jackman, T. (2014) Multiple origins of parthenogenesis, and a revised species phylogeny for the Southeast Asian butterfly lizards, Leiolepis. Biological Journal of the Linnean Society 113, 1080–1093. https://doi.org/10.1111/bij.12367
Chan, K.O., Brown, R.M., Lim, K.K.P., Ahmad, N. & Grismer, L.L. (2014) A new species of frog (Amphibia: Anura: Ranidae) of the Hylarana signata Complex from Peninsular Malaysia. Herpetologica 70, 228–240. https://doi.org/10.1655/HERPETOLOGICA-D-13-00057
Chan, K.O., Grismer, L.L. & Brown, R.M. (2014) Reappraisal of the Javanese Bullfrog complex, Kaloula baleata(Müller, 1836) (Amphibia: Anura: Microhylidae), reveals a new species from Peninsular Malaysia. Zootaxa 3900, 569–580. https://doi.org/10.11646/zootaxa.3900.4.7
Chan, K.O., Wood, P.L.J., Anuar, S., Muin, M.A., Quah, E.S.H., Sumarli, A.X.Y. & Grismer, L.L. (2014) A new species of upland Stream Toad of the genus Ansonia Stoliczka, 1870 (Anura: Bufonidae) from northeastern Peninsular Malaysia. Zootaxa 3764, 427–440. https://doi.org/http://dx.doi.org/10.11646/zootaxa.3764.4.3
Grismer, L.L., Belabut, D.M., Quah, E.S.H., Chan, K.O., Wood, P.L. & Hasim, R. (2014) A new species of karst forest-adapted Bent-toed Gecko (genus Cyrtodactylus Gray, 1827) belonging to the C. sworderi complex from a threatened karst forest in Perak, Peninsular Malaysia. Zootaxa 3755, 434–446. https://doi.org/10.11646/zootaxa.3755.5.3
Grismer, L.L., Wood, P.L., Anuar, S., Quah, E.S.H., Muin, M.A., Mohamed, M., Chan, K.O., Sumarli, A.X., Loredo, A.I. & Heinz, H.M. (2014) The phylogenetic relationships of three new species of the Cyrtodactylus pulchelluscomplex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia. Zootaxa3786, 359–381. https://doi.org/10.11646/zootaxa.3786.3.6
Grismer, L.L., Wood, P.L., Anuar, S., Riyanto, A., Ahmad, N., Muin, M.A., Sumontha, M., Grismer, J.L., Chan, K.O., Quah, E.S.H. & Pauwels, O.S.A. (2014) Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880, 1–147.
Grismer, L.L., Wood, P.L., Chan, K.O., Anuar, S. & Muin, M.A. (2014) Cyrts in the city: a new Bent-toed gecko (genus Cyrtodactylus) is the only endemic species of vertebrate from Batu Caves, Selangor, Peninsular Malaysia. Zootaxa 3774, 381–394. https://doi.org/10.11646/zootaxa.3774.4.6
Nurulhuda, Z., Senawi, J., Musa, F.H., Belabut, D., Chan, K.O., Md. Nor, S., Ahmad, N. & Chan, K.O. (2014) Species composition of amphibians and reptiles in Krau wildlife reserve, Pahang, Peninsular Malaysia. Check List 10, 335–343. https://doi.org/10.15560/10.2.335
2013
Chan, K.O., Blackburn, D.C., Murphy, R.W., Stuart, B.L., Emmett, D.A., Ho, C.T. & Brown, R.M. (2013) A new species of Narrow-mouthed frog of the genus Kaloula from Eastern Indochina. Herpetologica 69, 329–341. https://doi.org/10.1655/HERPETOLOGICA-D-12-00094
Grismer, L.L., Wood, P.L., Mohamed, M., Chan, K.O., Heinz, H.M., Sumarli, A.S.-I., Chan, J.A. & Loredo, A.I. (2013) A new species of karst-adapted Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) from a threatened karst region in Pahang, Peninsular Malaysia. Zootaxa 3746, 463–472. https://doi.org/10.11646/zootaxa.3746.3.5
2012
Johnson, C.B., Quah, E.S.H.S.H., Anuar, S., Muin, M.A., Wood, P.L.J., Grismer, J.L., Greer, L.F., Chan, K.O., Ahmad, N., Bauer, A.M. & Grismer, L.L. (2012) Phylogeography, geographic variation, and taxonomy of the Bent-toed gecko Cyrtodactylus quadrivirgatus Taylor, 1962 from Peninsular Malaysia with the description of a new swamp-dwelling species. Zootaxa 3406, 39–58.
2011
Ahmad, N., Farah, A.D., Chan, K.O., Belabut, D. & Muin, M.A. (2011) An update of herpetofaunal records from Bukit Fraser, Pahang, Peninsular Malaysia. Malaysian Applied Biology 40, 9–17.
Chan, K.O., Grismer, L.L., Anuar, S., Quah, E.S.H., Grismer, J.L., Wood, P.L., Muin, M.A. & Ahmad, N. (2011) A new species of Chiromantis Peters 1854 (Anura: rhacophoridae) from Perlis state park in extreme northern Peninsular Malaysia with additional herpetofaunal records for the park. Russian Journal of HerpetologyJournal of Herpetology 18, 253–259.
Chan, K.O., Grismer, L.L. & Grismer, J.L. (2011) A new insular, endemic frog of the genus Kalophrynus Tschudi, 1838 (Anura: Microhylidae) from Tioman Island, Pahang, Peninsular Malaysia. Zootaxa 68, 60–68.
Grismer, L.L., Grismer, J.L., Wood, P.L., Ngo, V.T., Neang, T. & Chan, K.O. (2011) Herpetology on the fringes of the Sunda shelf: a discussion of discovery, taxonomy, and tiogeography. Bonner Zoologische Monographien 57, 57–97.
Grismer, L.L., Quah, E.S.H., Siler, C.D., Chan, K.O., Wood, P.L., Grismer, J.L., Anuar, S. & Ahmad, N. (2011) Peninsular Malaysia’s first limbless lizard: a new species of skink of the genus Larutia (Böhme) from Pulau Pinang with a phylogeny of the genus. Zootaxa 2799, 29–40.
Ngo, V.T. & Chan, K.O. (2011) A new karstic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from Northern Vietnam. Zootaxa 3125, 51–63.
Norhayati, A., Chan, K.O., Daicus, B., Samat, A., Grismer, L.L. & Mohd Izzuddin, A. (2011) Potential biosites of significant importance in Langkawi Geopark: Terrestrial vertebrate fauna. Planning Malaysia 9.
Quah, E.S.H., Anuar, S.M.S., Grismer, L.L., Muin, M.A., Chan, K.O. & Grismer, J.L. (2011) Preliminary checklist of the herpetofauna of Jerejak Island, Penang, Malaysia. Malayan Nature Journal 63, 595–600.
van Rooijen, J., Chan, K.O., Grismer, L.L. & Ahmad, N. (2011) Estimating the herpetofaunal species richness of Pangkor Island, Peninsular Malaysia. Bonn zoological Bulletin 60, 3–8.
Savage, A.E., Grismer, L.L., Anuar, S., Chan, K.O., Grismer, J.L., Quah, E.S.H., Muin, M.A., Ahmad, N., Lenker, M. & Zamudio, K.R. (2011) First record of Batrachochytrium dendrobatidis infecting four frog families from Peninsular Malaysia. EcoHealth 8, 121–128. https://doi.org/10.1007/s10393-011-0685-y
2010
Chan, K.O. & Ahmad, N. (2010) A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389, 47–56.
Chan, K.O., Belabut, D. & Ahmad, N. (2010) A revised checklist of the amphibians of Peninsular Malaysia. Russian Journal of Herpetology 17, 202–206.
Chan, K.O. & Grismer, L.L. (2010) Re-assessment of the Reinwardt’s gliding frog, Rhacophorus reinwardtii (Schlegel 1840) (Anura: Rhacophoridae) in southern Thailand and Peninsular Malaysia and its re-description as a new species. Zootaxa 2505, 40–50.
Chan, K.O., Grismer, L.L., Anuar, S., Quah, E.S.H., Muin, M.A., Savage, A.E., Grismer, J.L., Ahmad, N., Remigio, A.C. & Greer, L.F. (2010) A new endemic rock gecko Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia. Zootaxa 68, 59–68.
Chan, K.O., van Rooijen, J., Grismer, L.L., Belabut, D., Muin, M.A., Jamaludin, H., Gregory, R. & Norhayati, A. (2010) First report on the herpetofauna of Pulau Pangkor, Perak, Malaysia. Russian Journal of Herpetology 17, 139–146.
Grafe, T.U., Norhayati, A., Chan, K.O., Daicus, B. & Senawi, J. (2010) A checklist of the Herpetofauna of Lanjak Entimau Wildlife Sanctuary, Sarawak Lanjak Entimau Wildlife Sanctuary “Hidden Jewel of Sarawak.” Akademi Sains Malaysia & Sarawak Forestry Department, Kuala Lumpur.
Grismer, L.L., Anuar, S., Quah, E.S.H., Muin, M.A., Chan, K.O., Grismer, J.L. & Ahmad, N. (2010a) A new spiny, prehensile-tailed species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia with a preliminary hypothesis of relationships based on morphology. Zootaxa 2625, 40–52.
Grismer, L.L. & Chan, K.O. (2010) Another new Rock Gecko (genus Cnemaspis strauch 1887) from Pulau Langkawi, Kedah, Peninsular Malaysia. Zootaxa 2419, 51–62.
Grismer, L.L., Chan, K.O., Grismer, J.L., Wood, P.L., Norhayati, A. & Ahmad, N. (2010) A checklist of the herpetofauna of the Banjaran Bintang, Peninsular Malaysia. Russian Journal of Herpetology 17, 147–160.
Grismer, L.L., Chan, K.O., Quah, E.S.H., Muin, M.A., Savage, A.E., Grismer, J.L., Ahmad, N., Greer, L.F. & Remegio, A.C. (2010) Another new, diminutive Rock Gecko (Cnemaspis Strauch) from Peninsular Malaysia and a discussion of resource partitioning in sympatric species pairs. Zootaxa 66, 55–66.
Lim, B.L., Wira, N.A., Chan, K.O., Daicus, B. & Norhayati, A. (2010) An updated checklist of the herpetofauna of Pulau Singa Besar, Langkawi , Peninsular Malaysia. Malaysian Applied Biology 39, 13–23.
Ngo, V.T. & Chan, K.O. (2010) A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa province, Vietnam. Zootaxa 2504, 47–60.
Thy, N., Grismer, L.L., Chan, K.O., Grismer, J.L., Wood. P. L. Jr. & Youmans, T.M. (2010) First report on the herpetofauna of Dalai Mountain in Phnom Samkos Wildlife Sanctuary, southwestern Cardamom Mountains, Cambodia. Cambodian Journal of Natural History, 127–143.
2009
Chan, K.O. (2009) The rediscovery of Enhydris pahangensis Tweedie, 1946. Hamadryad 34, 174–175.
Chan, K.O. & Ahmad, N. (2009) Distribution and natural history notes on some poorly known frogs and snakes from Peninsular Malaysia. Herpetological Review 40, 294–301.
Chan, K.O., Azman, M.S., Azlin, N. & Pan, K.A. (2009) Additions to the herpetofauna of Pasoh forest reserve, Negeri Sembilan, Peninsular Malaysia. Tropical Life Sciences Research 20, 71–80. https://doi.org/10.5281/zenodo.184144
Chan, K.O., Grismer, L.L., Ahmad, N. & Belabut, D. (2009) A new species of Gastrophrynoides (Anura: Microhylidae): an addition to a previously monotypic genus and a new genus for Peninsular Malaysia. Zootaxa68, 63–68.
Chan, K.O., Grismer, L.L., Sharma, D.S., Daicus, B. & Norhayati, A. (2009) New herpetofaunal records for Perlis state park and adjacent areas. Malayan Nature Journal 61, 277–284.
Chan, K.O., Grismer, L.L., Wood, P.L., Grismer, J.L. & Ahmad, N. (2009) Preliminary checklist of the herpetofauna of Pulau Besar, Melaka, Malaysia. Tropical Life Sciences Research 20, 81–87.
Grismer, L.L., Ahmad, N. & Chan, K.O. (2009) new faces from ancient Places: Uncovering Peninsular Malaysia’s Hidden lizard Diversity Part i: Geckos. Reptiles & Amphibians 16, 256–260.
Grismer, L.L., Ahmad, N. & Chan, K.O. (2009) A new, diminutive, upland Sphenomorphus Fitzinger 1843 (Squamata: Scincidae) from the Belum-Temengor forest complex, Peninsular Malaysia. Zootaxa 2312, 27–38.
Grismer, L.L., Ahmad, N., Chan, K.O., Belabut, D., Muin, M.A., Wood, P.L. & Grismer, J.L. (2009) Two new diminutive species of Cnemaspis strauch 1887 (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 2019, 40–56. https://doi.org/10.5281/zenodo.186045
Grismer, L.L. & Chan, K.O. (2009) A new species of karst-dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Sarawak, Borneo. Zootaxa 2246, 21–31.
Wood, P.L.J., Grismer, J.L., Grismer, L.L., Ahmad, N., Chan, K.O. & Bauer, A.M. (2009) Two new montane species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Peninsular Malaysia. Zootaxa 2012, 28–46. (doi:http://dx.doi.org/10.11646/zootaxa.2012.1.2)
2008
Chan, K.O., Norhayati, A. & Senawi, J. (2008) Kekayaan spesies herpetofauna di Kuala Gandah menggunakan kaedah perangkap lubang berpagar. In: E. Sivananthan, Y. Siti Hawa, A. H. Abd Kadir, H. Hasdi, B. L. Lim, H. Rosli, A. Noorlidah, M. N. Shukor, and I. Rahmah (Eds), Biodiversity Resources and Conservation Status in Peninsular Malaysia. Department of Wildlife & National Parks, pp. 71–84.
Chan, K.O. & Grismer, L.L. (2008) A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Selangor, Peninsular Malaysia. Zootaxa 57, 49–58.
Norhayati, A., Rahman, M.R., Senawi, J., Belabut, D. & Chan, K.O. (2008) A checklist of amphibians at Taman Negara Kelantan National Park, Kuala Koh, Peninsular Malaysia. Journal of Wildlife and Parks 25: 1–7.
Grismer, L.L. & Chan, K.O. (2008) A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. Zootaxa 1771, 1–15.
Grismer, L.L., Chan, K.O., Grismer, J.L., Wood, P.L.J. & Belabut, D. (2008) Three new species of Cyrtodactylus(Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 1921, 1–23.
Grismer, L.L., Chan, K.O., Nasir, N. & Sumontha, M. (2008) A new species of karst dwelling gecko (genus Cnemaspis Strauch 1887) from the border region of Thailand and Peninsular Malaysia. Zootaxa 68, 51–68.
Grismer, L.L., Grismer, J.L., Wood, P.L. & Chan, K.O. (2008) The distribution, taxonomy, and redescription of the geckos Cnemaspis affinis (Stoliczka 1887) and C. flavolineata (Nicholls 1949) with descriptions of a new montane species and two new lowland, karst- dwelling species from Peninsular Malays. Zootaxa 1931, 1–24. https://doi.org/http://dx.doi.org/10.11646/zootaxa.1931.1.1
2007
Norhayati, A., Senawi, J., Daicus, B. & Chan, K.O. (2007) Amphibian diversity of Taman Rimba Kenong. In: A. Muda, A. A. Rahman, C. H. Hassan, H. L. Koh, A. Norhayati, and A. Latiff (Eds), Pengurusan, Persekitaran Fizikal dan Kepelbagaian Biologi. Forestry Department of Peninsular Malaysia, Kuala Lumpur.